I completed AC1 a while back and it took no time flat, AC1's lore isn't exactly hard to understand because you are just Desmond trapped in Abstergo. The onk pieces of lore in the game are the Apple of Eden and The Eagle's Vision in Desmond's Room. Blood splattered on the walls from the previous Test Subject, Subject 16. That's all you know of 16.
You find out in ACII that Subject 16 is a much bigger part than what you probably first thought. You find The Truth through his multiple visions he left around Ezio's Italy for you to find. Outlining the betrayal, hatred, and pettiness of War, Religion, and Love. The ending, The Truth, reveals that Adam and Eve were that of, or slaves of, a much more advanced race than what was previously thought, outlining The First Civilization, Juno, Minerva, and Jupiter in particular who are the outlines of the future. Peering through their godforsaken world into the ruined and rebuilt future, seeing what would once happen again, and warning future civilizations of the coming apocalypse. They discovered that their one and only hope would lie an uncountable am ou until of time later in Desmond. The end of ACII, spoilers, sees Minerva conversing with Ezio only to turn, stop, turn to the camera mid sentence, and whisper "Desmond?', much to Ezio's confusion. This would be Desmond's first contact with The First Civilization.
In Assassin's Creed Brotherhood, you once again are searching for Subject 16's truth, only this time it is the truth of a betrayal as a being of computer algorithm stood before Desmond and told him of the plan to sabotage, not only the Assassin's, but also the world. The betrayal of the one person that Desmond cares for most, Lucy Stillmam. He learns that Lucy all along was working for the Templars and sought to use Desmond to further their plan. I'm the ending to Brotherhood Juno takes matters into his own hands and after Desmond, Lucy, and the others find the Apple of Eden beneath the Roman Colosseum, Juno takes control of Desmond, forcing him to stab Lucy and ending her plot. This sends Desmond into a coma, the same coma that trapped Subject 16 as well.
In Revelations there is no Truth. There are no lies being told for Desmond to be informed of. Revelations is as much as a duality between Subject 16, Clay Kaczmarek, and Subject 17, Desmond Miles, as it is between Altaïr and Ezio. But you would never know that from the base game. Despite Clay showing up and bantering with Desmond explaining to him Animus Island and the path that he has to traverse to escape his nightmare, Clay really has very little to do with the story. In the end of Ezio's final game he meets with Jupiter who tells Ezio and Desmond the signs, the apocalypse. And the story ends.
In the Lost Archive though we discover Clay's story. As he, like Altaïr and Ezio, is a vessel to assist Desmond. His story is quite sad. As he is dealing with his own family turmoil. Wanting to be a history teacher but being denied by his father in respect of being an Engineer to only be in turn told that the Engineering field, Computer Engineering, was unacceptable. He was told, "Kaczmarek's don't sit behind computers, they build with their hands". All of this turmoil while still fulfilling his training and duty as an Assassin. Finally the day came when he was to infiltrate Abstergo, the Corporate front for the Templars, and enter the Animus. This only made his family relations worse. He had finally found his purpose, but all his father was worried about was that he didn't receive a check from Clay that month. The Animus tears him apart. Unrelenting stress and demands from Abstergo pushed him to his breaking point. He finally built enough money from the tests to send a check to his family. He receives an email around this time from his drunken and drugged mother. She was leaving. And soon after discovered that Lucy, his closest friend and his only hope of survival, was a traitor. She found out and told him that she promised she would protect him and would continue to do so but he could never leave. In the end Clay's final moments were spent inside the Animus, informed by Juno, trying to make things right. Trying to warn a man he'd never met. Trying to save the world. Stuck within a loop of his own life reliving the time from when he received his mission to his breaking point of him falling through the clouds again and again until he finally broke the loop, escaped the animus and did what he had to do and took his own life for the sole purpose of showing Desmond The Truth.